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Neurotransmission at chemical synapses is restricted to specialized locations with the presynaptic plasma membrane known as active zones (AZ). There, a tight network of multi-domain scaffolding proteins, the cytomatrix in the AZ (CAZ), orchestrates the controlled exoand endocytosis of synaptic vesicles in space and time. CAZ components like Bassoon (Bsn), Piccolo/Aczonin (Pclo), RIM, ELKS/CAST, and Munc13 contribute to synaptic transmission either by straight participating in vesicle priming, docking, and retrieval, or by supplying interaction internet sites for molecules involved in these processes [1,2]. Morphological variations on the AZ will be the ribbon synapses of sensory neurons of the visual and auditory systems [3]. Whereas the CAZ at traditional chemical synapses is often a additional or less two-dimensional specialization, ribbon synapses harbor a three-d.